@articleDe Jaegher_Hoyer_2016, title=By-product mutualism and the ambiguous effects of harsher environments – A game-theoretic model, volume=393, DOI=10.1016/j.jtbi.2015.12.034, journal=Journal of Theoretical Biology, publisher=Elsevier BV, author=De Jaegher, Kris and Hoyer, Britta, year=2016, pages=82–97